We are looking for an exceptional candidate to carry out a combined theoretical and experimental PhD. The student will receive training from two labs: Professor Peter Latham (Gatsby Computational Neuroscience Unit; theory) and Professor Michael Hausser (Wolfson Institute for Biomedical Research; experiment). The candidate will initially be based at the Gatsby Unit with Professor Latham, taking courses in computational neuroscience and developing theories at the circuit level; the rest of the PhD will be with Professor Hausser testing those theories using advanced experimental tools for linking neural circuit function with behavior (most notably targeted optogenetics  and fast 2-photon imaging). The applicant should have a strong analytical background and a keen interest in neuroscience. Experience in an experimental lab is desirable but not absolutely necessary. The applicant will be fully funded for four years. Start date is flexible (but ideally by 1 October 2014).  Closing date is May 15! Applicants should apply directly to Professor Latham (pel@gatsby.ucl.ac.uk). Please send, in PDF or plain text format where possible, the following: - CV - statement of research interests, - transcript(s) for previous degrees, and arrange for three academic letters of recommendation, also to pel@gatsby.ucl.ac.uk . For further details of research interests please visit our websites: Latham:  http://www.gatsby.ucl.ac.uk/~pel/ Hausser: http://www.dendrites.org/